diff --git a/app/models/query.rb b/app/models/query.rb
old mode 100644
new mode 100755
index b0b39da..84d5b20
--- a/app/models/query.rb
+++ b/app/models/query.rb
@@ -235,7 +235,7 @@ class Query < ActiveRecord::Base
     :list => [ "=", "!" ],
     :list_status => [ "o", "=", "!", "c", "*" ],
     :list_optional => [ "=", "!", "!*", "*" ],
-    :list_subprojects => [ "*", "!*", "=" ],
+    :list_subprojects => [ "*", "!*", "=", "!" ],
     :date => [ "=", ">=", "<=", "><", "<t+", ">t+", "><t+", "t+", "t", "ld", "w", "lw", "l2w", "m", "lm", "y", ">t-", "<t-", "><t-", "t-", "!*", "*" ],
     :date_past => [ "=", ">=", "<=", "><", ">t-", "<t-", "><t-", "t-", "t", "ld", "w", "lw", "l2w", "m", "lm", "y", "!*", "*" ],
     :string => [ "=", "~", "!", "!~", "!*", "*" ],
@@ -661,12 +661,19 @@ class Query < ActiveRecord::Base
 
   def project_statement
     project_clauses = []
-    if project && !project.descendants.active.empty?
+    active_subprojects_ids = []
+
+    active_subprojects_ids = project.descendants.active.map(&:id) if project
+    if active_subprojects_ids.any?
       if has_filter?("subproject_id")
         case operator_for("subproject_id")
         when '='
           # include the selected subprojects
-          ids = [project.id] + values_for("subproject_id").each(&:to_i)
+          ids = [project.id] + values_for("subproject_id").map(&:to_i)
+          project_clauses << "#{Project.table_name}.id IN (%s)" % ids.join(',')
+        when '!'
+          # exclude the selected subprojects
+          ids = [project.id] + active_subprojects_ids - values_for("subproject_id").map(&:to_i)
           project_clauses << "#{Project.table_name}.id IN (%s)" % ids.join(',')
         when '!*'
           # main project only
diff --git a/test/unit/query_test.rb b/test/unit/query_test.rb
old mode 100644
new mode 100755
index 6d27106..6db72d6
--- a/test/unit/query_test.rb
+++ b/test/unit/query_test.rb
@@ -682,15 +682,15 @@ class QueryTest < ActiveSupport::TestCase
     Member.create!(:project_id => 1, :principal => other_group, :role_ids => [1])
     User.current = user
 
-    with_settings :issue_group_assignment => '1' do 
+    with_settings :issue_group_assignment => '1' do
       i1 = Issue.generate!(:project_id => 1, :tracker_id => 1, :assigned_to => user)
       i2 = Issue.generate!(:project_id => 1, :tracker_id => 1, :assigned_to => group)
       i3 = Issue.generate!(:project_id => 1, :tracker_id => 1, :assigned_to => other_group)
-  
+
       query = IssueQuery.new(:name => '_', :filters => { 'assigned_to_id' => {:operator => '=', :values => ['me']}})
       result = query.issues
       assert_equal Issue.visible.where(:assigned_to_id => ([2] + user.reload.group_ids)).sort_by(&:id), result.sort_by(&:id)
-  
+
       assert result.include?(i1)
       assert result.include?(i2)
       assert !result.include?(i3)
@@ -1810,4 +1810,21 @@ class QueryTest < ActiveSupport::TestCase
     ActiveRecord::Base.default_timezone = :local # restore Redmine default
   end
 
+  def test_filter_on_subprojects
+    query = IssueQuery.new(:name => '_', :project => Project.find(1))
+    filter_name = "subproject_id"
+    assert_include filter_name, query.available_filters.keys
+
+    # "is" operator should include issues of parent project + issues of the selected subproject
+    query.filters = {filter_name => {:operator => '=', :values => ['3']}}
+    issues = find_issues_with_query(query)
+    assert_equal [1, 2, 3, 5, 7, 8, 11, 12, 13, 14], issues.map(&:id).sort
+
+    # "is not" operator should include issues of parent project + issues of all active subprojects - issues of the selected subprojects
+    query = IssueQuery.new(:name => '_', :project => Project.find(1))
+    query.filters = {filter_name => {:operator => '!', :values => ['3']}}
+    issues = find_issues_with_query(query)
+    assert_equal [1, 2, 3, 6, 7, 8, 9, 10, 11, 12], issues.map(&:id).sort
+  end
+
 end